Teams that played in the FA Women's National League Division One were given exemption to the second round qualifying, while teams in the Northern and Southern Premier Divisions entered in the second round proper.
The second round qualifying saw one tie cancelled due to the withdrawal of St Nicholas.
[3] It was added by The Football Association as a result of increased entries into the competition.
[4] Fifty three matches were scheduled for the preliminary round, to be played by Sunday 26 August 2018.
Matches were played on the scheduled date of Sunday 2 September 2018, except two delays caused by matches from the previous round not yet having taken place and Swindon Spitfires v Southampton FC Women delayed due to the removal of Moneyfields from the competition.
[11] Most matches were played on Sunday 11 November 2018, the only exception being New London Lionesses v AFC Wimbledon which were postponed from the original scheduled date to the following week due to a waterlogged pitch.
[16] The 40 teams taking part consisted of 25 FA Women's National League Northern and Southern Division teams exempted to this stage, plus the 15 match winners from the previous round.
[18] The 32 teams taking part consists of 22 FA Women's Super League and FA Women's Championship teams exempted to this stage, plus the ten match winners from the previous round.