The 49th Reserve Division initially fought on the Eastern Front, entering the line in mid-October.
It fought in the Battle of Łódź in November–December 1914 and then spent most of the period until mid-1915 fighting along the Rawka and Bzura Rivers.
In July and August 1915 it fought in the Battle of Warsaw and then participated in the follow-on engagements as German forces drove the Russians back to a line along the Shchara and Servech Rivers.
The division then fought in various parts of the line, including in the Carpathians, until January 1917, when it was transferred to the Western Front.
Over the course of the war, other changes took place, including the formation of artillery and signals commands and the enlargement of combat engineer support to a full pioneer battalion.