Part of the 1st Army Tank Brigade, 7th RTR saw service in France in May 1940, alongside the 4th Royal Tank Regiment and the 50th (Northumbrian) Infantry Division during the Battle of Dunkirk and was soon evacuated at Dunkirk, and abandoning most of their vehicles.
The regiment attacked and destroyed the Italian "Maletti Group" at the Nibeiwa Camp.
The 7th RTR also supported the attack of the 11th Brigade on Italian positions at the Tummar camps.
The new 7th RTR joined the 31st Tank Brigade in Normandy on 18 June 1944 and subsequently took part in heavy fighting in Operation Epsom.
It participated in the capture of Le Havre and, while detached from its Brigade, it played an active part in the early days of the containment of Dunkirk.