Accipitridae

The Accipitridae have a cosmopolitan distribution, being found on all the world's continents (except Antarctica) and a number of oceanic island groups.

The genus level cladogram of the Accipiridae shown below is based on a densely sampled molecular phylogenetic study of the Accipitridae by Therese Catanach and collaborators that was published in 2024.

[7] The number of species in each genus is based on the list maintained by Frank Gill, Pamela C. Rasmussen and David Donsker on behalf of the International Ornithological Committee (IOC).

As evidenced by fossils like Pengana, some 25 mya, accipitrids in all likelihood rapidly acquired a global distribution, initially probably extending even to Antarctica.

Specimen AMNH FR 2941, a left coracoid from the Late Eocene Irdin Manha Formation of Chimney Butte (Inner Mongolia), was initially assessed as a basal mid-sized buteonine;[15] it is today considered to be more likely to belong in the Gruiformes genus Eogrus.

Until the 14th century, even these huge vultures were surpassed by the extinct Haast's eagle (Hieraaetus moorei) of New Zealand, which is estimated to have measured up to 140 cm (55 in) and to have weighed 15 to 16.5 kg (33 to 36 lb) in the largest females.

In a majority of species, such as generalist hunters and rodent-, reptile-, fish-, and insect-hunting specialists, the dimorphism is less, usually between a 5% to 30% size difference.

In the carrion-eating Old World vultures and snail eating kites, the difference is largely non-existent, though sometimes the female may average slightly heavier.

The plumage of the Accipitridae can be striking, but rarely utilises bright colours; most birds use combinations of white, grey, buff, brown and black.

[25] The senses of the Accipitridae are adapted to hunting (or scavenging), and in particular their vision is exceptional, with some large accipitrids such as the wedge-tailed eagle and Old World vultures having over twice the visual acuity of a typical human.

Prey is usually captured and killed in the powerful talons of the raptor and then carried off to be torn apart with a hooked bill for eating or feeding to nestlings.

However, most have a preference for a certain type of prey, which in harriers and the numerous buteonine hawks (including more than 30 species in the genus Buteo) tends towards small mammals such as rodents.

Among the raptors that mainly favor small mammals, harriers generally hunt by hovering over openings until they detect their prey and descend upon them.

[30] Buteonine hawks usually watch for prey from a perch but most species will also readily hunt on the wing, including from a high soar.

[23] The diet of the honey-buzzards includes not only the adults and young of social insects such as wasps and bees, but the honey and combs from their nests.

However, many accipitrids of almost all sizes have been recorded as capturing and then flying with prey of equal weight or even slightly heavier than themselves in their talons, a feat that requires great strength.

This has the advantage of providing a surplus of food but has the disadvantage of potentially attracting scavengers or other predators which can steal the kill or even attack the feeding accipitrid.

Bazas and forest hawks in the genus Accipiter may take reptiles from trees whilst other species may hunt them on the ground.

[37][38] In terms of their reproductive biology and socio-sexual behavior, accipitrids share many characteristics with other extant groups of birds that appear not be directly related, but all of which have evolved to become active predators of other warm-blooded creatures.

Some of the characteristics shared with these other groups, including falcons, owls, skuas and shrikes, are sexual dimorphism in size, with the female typically larger than the male; extreme devotion of breeding pairs to each other or to a dedicated nesting site; strict and often ferocious territorial behavior; and, on hatching, occasional competition amongst nestlings, including regular siblicide in several species.

While a single devoted breeding pair is considered typical, research has revealed that in varied accipitrids, multiple birds engaging in nesting behavior is more commonly than previously thought.

Some harriers have evolved to become polygynous, with a single smaller male breeding with and then helping multiple females raise young.

Nest sites are typically in relatively secure places, such as the crook of a large tree or an ample cliff ledge, and can vary in elevation from the flat ground of prairies or steppe to near the peaks of the tallest mountains.

Once independent of their parents, young accipitrids often most wander for considerable stretches of time, ranging from 1 to 5 years, before they attain maturity.

Most accipitrids have distinct plumages in their immature stage, which presumably serves as a visual cue to others of their species and may allow them to avoid territorial fights.

Shortly after attaining mature plumages, pairs form, with a male typically displaying, often in flight but sometimes vocally, to win over a female.

Neophrontops americanus fossil
Neogyps errans fossil
Portrait of a subadult bald eagle , showing its strongly hooked beak and the cere covering the base of the beak.
The palm-nut vulture is an unusual frugivorous accipitrid, but will also consume fish, particularly dead fish
Oriental honey-buzzard Pernis ptilorhyncus