[3] Recent phylogenetic studies, however, conclude that morphological, geographical, and genetic evidence make it clear that it is a separate species.

[6] Like other members of the genus, the butterfly is named "sister" for its black and white markings on the forewing that resemble a nun's habit.

[4][6] They also differ morphologically from A. eulalia in usually being smaller and in the presence of a second orange bar in the upper proximal corner of the hindwings.

[2][4][9] Other species which resemble A. californica (though not as closely) includes other members of the genus Adelpha, and the Lorquin's admiral (Limenitis lorquini).

They are laid singly on tips of leaves of oak plants (genus Quercus), on the upper surface.

After eight days it molts again into the third instar which is a pale brownish yellow covered densely in yellowish conical tubercles.

It molts after nine days into the last instar which is olive green with brown undersides and legs, it is densely covered with yellowish tubercles (particularly at the bottom surfaces).

[5] This diet makes A. californica unpalatable to predators, which might explain why so many other species have formed a mimicry complex around it.

[4] The adults are commonly found flying near the upper branches of oak trees or perching near small streams and canyons.

Both sexes also feed on nectar from flowers (though this is rare), as well as drink from rotting fruit, sap, and animal droppings.

Depending on seasonal conditions and elevation, the species produces one to three generations annually, with the adults usually flying from March through November.