Archaeamphora

Fossil material assigned to this taxon originates from the Yixian Formation of northeastern China, dated to the Early Cretaceous (around 143 to 101 million years ago).

[9] The generic name Archaeamphora is derived from the Greek αρχαίος, archaíos ("ancient"; combining form in Latin: archae-), and ἀμφορεύς, amphoreus ("pitcher").

The specific epithet longicervia is derived from the Latin longus ("long") and cervicarius ("with a neck"), in reference to the characteristic constriction in the pitcher-like structures of this species.

[1] The fossil material of A. longicervia was subjected to chemical analysis for oleanane, considered a key marker differentiating angiosperms from gymnosperms.

Of particular note is the similarity between the thick semi-circular structures on the lid of A. longicervia and the large nectar-secreting "bubble" present on the upper posterior portion of Heliamphora exappendiculata[nb 1] pitchers.

This variety differs from the type material of A. longicervia in having pitchers that lack any constriction before the mouth, instead gradually expanding from the petiole into a hollow trumpet-like shape.

[...] Another contradiction is that except for Archaeamphora, there is no other evidence to suggest that Sarraceniaceae evolved outside the New World, to which all extant members of the family are endemic.Wong et al. (2015) put forward a new perspective as follows:[9] Archaeamphora longicervia H. Q. Li was described as an herbaceous, Sarraceniaceae-like pitcher plant from the mid Early Cretaceous Yixian Formation of Liaoning Province, northeastern China.

We demonstrate that putative pitchers of Archaeamphora are insect-induced leaf galls that consist of three components: (1) an innermost larval chamber; (2) an intermediate zone of nutritive tissue; and (3) an outermost wall of sclerenchyma.

Archaeamphora is not a carnivorous, Sarraceniaceae-like angiosperm, but represents insect-galled leaves of the previously reported gymnosperm Liaoningocladus boii G. Sun et al. from the Yixian Formation.The area inhabited by A. longicervia is thought to have experienced significant climatic fluctuations during the Early Cretaceous, ranging from arid or semi-arid to more humid conditions.

Artist's reconstruction of Archaeamphora longicervia as pitcher plant, after Li (2005) [ 1 ]