Biseridens

[2][3] Primarily understood from the recently discovered, most-well preserved specimen, Biseridens is most notably distinguishable as an anomodont due to its short snout, dorsally elevated zygomatic arch, and presence of a septomaxilla that distinctly lacks a facial exposure between the nasal and maxilla.

[1] Biseri- is Latin for "double rows"; -dens originated from Greek "odons" for "teeth" At a clade level, anomodonts were terrestrial vertebrates of all sizes that dominated the Late Permian.

[4] On the basis of converging faunal evidence and comparison to other regions, the Xidagou Formation has been considered to be Middle Permian[6] and home to the oldest therapsid fauna, suggesting that anomodonts had a Laurasian rather than a Gondwanan origin.

[8] The more recently discovered specimen includes a nearly complete skull (lacking the occiput) with lower jaws, found with a set of 14 articulated vertebrae.

This new specimen was preserved in mudrock and also discovered in the upper part of Xidagou Formation in Dashankau, Yumen in Gansu.

Additionally, the mandibular fenestra is absent from the lower jaw, the transverse flange of the pterygoid extends laterally, and there is contact between the tabular and opisthotic, or the posterior part of the otic capsule.

[8] Although relatively primitive, a number of shared traits (synapomorphies) ally Biseridens with anomodonts including the shortened snout, raised zygomatic arch and exclusion of the septomaxilla between the maxilla and nasals.

However, it retains a number of primitive traits that exclude it from the more derived anomodonts, including the differentiated tooth row, palatal teeth, contact between tabular and opisthotic; lateral process of transverse flange of pterygoid free of posterior ramus and absence of mandibular foramen.

[1] Biarmosuchia Gorgonopsia Dinocephalia Biseridens Anomocephalus Otsheria Ulemica Suminia Patranomodon Galeops Eodicynodon

Holotype skull of Biseridens qilianicus on display at the Paleozoological Museum of China.