This stands in contrast the strictly marine fossils found in the underlying Dzheirantui Formation, and indicates that the Bissekty was formed during the regression of a saltwater sea.
The coastline expanded inland again in the upper portion of the Bissekty, represented by a proportional increase of fully aquatic species, which were almost completely absent from the middle period of the formation.
Eventually the area was again completely underwater, during the time period represented by the later Aitym Formation, which preserves coastal marine sediments.
[2] The Bissekty Formation is notable for preserving the most abundant Turonian land animal fossils in Eurasia, and the most diverse fauna of Late Cretaceous eutherians (placental mammals and relatives) in the world.
[2] Listings and accompanying information are based on a survey of the Bissekty Formation published by Cory Redman and Lindsey Leighton in 2009 unless otherwise noted.