On the underside of the cap there is usually a spongy surface with pores, instead of the gills typical of mushrooms.
The micromorphology and molecular phylogeny of the order Boletales have established that it also contains many gilled, puffball, and other fruit body shapes.
Examples of these fungi include Chroogomphus, Gomphidius, Phylloporus, Paxillus, Tapinella, Hygrophoropsis,[2] and Scleroderma.
[3] The genus Boletus was originally broadly defined and described by Carl Linnaeus in 1753, containing all pored mushrooms.
[4] Since then, gradually other genera have been defined, such as Tylopilus by Karsten in 1881,[5] and old names such as Leccinum and Suillus resurrected or redefined.