Afterwards he served as an assistant to Charles Flahault (1884–1888) in Montpellier and to Philippe Van Tieghem (1885–1891) in Paris.
[2] In 1891 he received his doctorate in Paris with the thesis Sur les feuilles de quelques Monocotylédones aquatiques (On the leaves of some aquatic monocots).
[3] In 1892 he attained a professorship at the University of Lyon, later serving as a professor of botany at the Faculty of Sciences of Bordeaux (1901–1932).
[2] He is known for his investigations of Phaeophyceae, being a taxonomic authority of numerous brown algae species.
[8] He also made contributions to Narcisse Patouillard's Catalogue raisonné des plantes cellulaires de la Tunisie.