Crabronidae

Crabroninae Dinetinae The Crabronidae is a large family of wasps within the superfamily Apoidea.

Of these lineages of Apoidea, only three were not included within Crabronidae in the past: Ampulicidae, Sphecidae, and Anthophila.

The following phylogenetic tree is based on Sann et al., 2018, which used phylogenomics to demonstrate that both the bees (Anthophila) and the Sphecidae arose from within the former Crabronidae, which is therefore paraphyletic, and which they suggested should be split into several families; the former family Heterogynaidae nests within the Bembicidae, as here defined.

[3][4] Ampulicidae Astatidae Bembicidae Sphecidae (sensu stricto) Crabronidae (sensu stricto) Mellinidae Pemphredonidae Philanthidae Psenidae Ammoplanidae Anthophila (bees) Further analysis by Sann et al. in 2021 resulted in the former Entomosericini (from Pemphredoninae) and Eremiaspheciinae being elevated to family status as Entomosericidae and Eremiaspheciidae, respectively.

The true phylogentic placements for both small families remain unresolved but outside of either Pemphredonidae, Psenidae, and the lineage comprising Ammoplanidae and bees.