The formation consists of coarse conglomerates, sandstone, mudstone, and thin coal layers deposited in deltaic and lacustrine environments.
In a 2006 study, Gao & Ren criticized He et al. for not including enough specifics and detail in their paper, and also took issue with their radiometric dating of the Daohugou tuff.
The authors stated that"vertebrate fossils such as Liaoxitriton, Jeholopterus and feathered maniraptorans show much resemblance to those of the Yixian Formation.
[8] Later in 2006, Liu et al. published their own study of the age of the Daohugou beds, this time using Zircon Uranium-lead dating on the volcanic rocks overlying and underlying salamander-bearing layers (salamanders are often used as index fossils).
[1][9] A 2012 study by Gao and Shubin agreed with this assessment, and reported an Argon–argon date of 164 plus or minus 4 million years ago for the Daohugou horizon.