[52] He had previously (1994–1995) described Habromorula as a separate genus, containing 14 species that were earlier classified with Spinidrupa sensu lato.

[53] In 2004, he placed Azumamorula, Oppomorus, and Habromorula into Morula as subgenera, and moved the whole group from subfamily Rapaninae to Ergalataxinae.

[54] In 2012–2013, Claremont et al. published a study of sequences for four genes across numerous species of Morula and related molluscs.

M. ceylonica, M. granulata, M. marginalba, M. musiva, and an undescribed species formed a distinct clade related to Muricodrupa.

In response to the gene-sequence phylogeny, Houart, Zuccon and Puillandre formally described new genera in 2019: Claremontiella, Murichorda, and Lauta.