Some exhibit rare feeding habits among ground beetles, including both omnivorous and even herbivorous species.

[3] With a variety of behaviors and habitats, Harpalinae species are predicted to have had a diversification rate greater than that of the angiosperms.

[4] Fossil records and evolutionary history show that tribes within Harpalinae likely arose and diversified during the mid-Cretaceous period.

[1] The variety of morphologies within Harpalinae make phylogenetic placements of tribes harder to establish without the support of genetic evidence.

[2] Molecular phylogenies may even point to such a rapid speciation of Harpalinae, showing some of these tribes arising at the same time.