In 972, Hillesley was recorded as Hildeslei (meaning "clearing belonging to Hild"), a tything of the parish of Hawkesbury.

[4] Pevsner notes the following buildings: The local church is St Giles – it was designed in 1851 by the amateur, Rev.

The greyish-white limestone farmhouse known as "Lovettswood," a prominent landmark, takes its name from the Lyvet family, who were lords of the manor of Hillesley in the 12th and 13th centuries.

Village amenities include a primary school, the church, a playing field hosting cricket, a tennis court and club, allotments and a mother and toddler group.

Until recent years, the village field hosted a successful football team, competing in local leagues.