[1] Crown without bony ridges; snout short, truncated; interorbital space flat, as broad as the upper eyelid; tympanum very distinct, vertically oval, quite as large as the eye and close to it.
First finger a little longer than second; toes barely half webbed, with irregular spinose tubercles beneath, from which the so-called subarticular are hardly distinguishable; two small metatarsal tubercles; no tarsal fold.
Upper parts studded with round tubercles of various sizes; parotoids prominent, subcircular.
Grey-brown or olive above, with irregular dark brown spots, vertical bars on the upper lip, and cross bands on the limbs; lower surfaces dirty white, with darker spots; the male's throat brown.
Male with a subgular vocal sac and, during the nuptial period, black rugosities on the inner fingers.