Isopods have various feeding methods: some eat dead or decaying plant and animal matter, others are grazers or filter feeders, a few are predators, and some are internal or external parasites, mostly of fish.
[3][8] The dorsal (upper) surface of the animal is covered by a series of overlapping, articulated plates which give protection while also providing flexibility.
The endopods (inner branches of the pleopods) are modified into structures with thin, permeable cuticles (flexible outer coverings) which act as gills for gas exchange.
[3] Isopods belong to the larger group Peracarida, which are united by the presence of a special chamber under the thorax for brooding eggs.
[14] In the deep sea, members of the suborder Asellota predominate, to the near exclusion of all other isopods, having undergone a large adaptive radiation in that environment.
[14] The largest isopod is in the genus Bathynomus and some large species are fished commercially for human food in Mexico, Japan and Hawaii.
Nowadays, the members of this formerly widespread suborder form relic populations in freshwater environments in South Africa, India and Oceania, the greatest number of species being in Tasmania.
The more advanced long-tailed isopods are mostly endemic to the southern hemisphere and may have radiated on the ancient supercontinent of Gondwana soon after it broke away from Laurasia 200 million years ago.
The short-tailed forms may have been driven from the shallow seas in which they lived by increased predatory pressure from marine fish, their main predators.
can run rapidly on land and many terrestrial species can roll themselves into a ball when threatened, a feature that has evolved independently in different groups and also in the marine sphaeromatids.
[9][24][25] Isopods have a simple gut which lacks a midgut section; instead there are caeca connected to the back of the stomach in which absorption takes place.
The structure of the stomach varies, but in many species there is a dorsal groove into which indigestible material is channelled and a ventral part connected to the caeca where intracellular digestion and absorption take place.
[9] Isopods are detritivores, browsers, carnivores (including predators and scavengers), parasites, and filter feeders, and may occupy one or more of these feeding niches.
Limnoria lignorum, for example, bores into wood and additionally feeds on the mycelia of fungi attacking the timber, thus increasing the nitrogen in its diet.
The larvae of the Gnathiidae family and adult cymothoidids have piercing and sucking mouthparts and clawed limbs adapted for clinging onto their hosts.
Fertilisation only takes place when the eggs are shed soon after a moult, at which time a connection is established between the semen receptacle and the oviduct.
The lack of a swimming phase in the life cycle is a limiting factor in isopod dispersal, and may be responsible for the high levels of endemism in the order.
[31] Terrestrial isopods play an important role in many tropical and temperate ecosystems by aiding in the decomposition of plant material through mechanical and chemical means, and by enhancing the activity of microbes.
[33] Macro-detritivores, including terrestrial isopods, are absent from arctic and sub-arctic regions, but have the potential to expand their range with increased temperatures in high latitudes.
They are subject to evaporation, especially from their ventral area, and as they do not have a waxy cuticle, they need to conserve water, often living in a humid environment and sheltering under stones, bark, debris or leaf litter.
Moisture is achieved through food sources or by drinking, and some species can form their paired uropodal appendages into a tube and funnel water from dewdrops onto their pleopods.
Members of the families Ligiidae and Tylidae, commonly known as rock lice or sea slaters, are the least specialised of the woodlice for life on land.