Sir John Ernest Walker (born 7 January 1941) is a British chemist who won the Nobel Prize in Chemistry in 1997.

[11][12] Walker began his study of peptide antibiotics with Edward Abraham at Oxford in 1965 and received his Doctor of Philosophy degree in 1969.

This resulted in an invitation to work at the Laboratory of Molecular Biology of the Medical Research Council, which became a long-term appointment.

Scientists trained in Walker's group at the MRC Laboratory of Molecular Biology in Cambridge or MRC Mitochondrial Biology Unit have gone on to determine crystal bacterial complex I and cryo-EM maps of mitochondrial complex I and vacuolar type ATPases.

[14] He shared his Nobel Prize with the American chemist Paul D. Boyer for their elucidation of the enzymatic mechanism underlying the synthesis of adenosine triphosphate.

[2] Walker is also a Foreign Associate of the National Academy of Sciences and an Honorary Fellow of St Catherine's College, Oxford.

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