The long-beaked fungus gnats, formerly placed in a separate family Lygistorrhinidae, have been placed into Keroplatidae as subfamily Lygistorrhininae.

[1] They are generally forest dwellers found in the damp habitats favoured by their host fungi.

Larvae both feed on fungi and are predatory - they can spin webs by secreting acid fluids, which they use to kill smaller invertebrates and capture spores.

The fossil record of the family extends back into the Cretaceous, with the oldest named member, Lebanognoriste known from the Barremian aged Lebanese amber, other Cretaceous species are known from the Spanish, Burmese, Bezonnais, Taimyr and Canadian ambers.

Despite the similarities in function and appearance, the bioluminescent systems of the three genera are not homologous and are believed to have evolved separately.