The abundant fossil remains were deposited by rivers and estuaries crossing an arid to semi-arid environment, between several marine transgressions.

Fossils from the Kuldana Formation have been studied since long before the name was established, with some of the earliest papers being those of Pilgrim, published in 1940.

Gingerich did however come to revise his 1983 age estimate in a paper published in 2003 that dealt with the stratigraphy observed across different localities in order to obtain a more reliable result.

At Banda Daud Shah, Gingerich recovered a late Early Eocene to early Middle Eocene (P9 or P10) age based on the planktonic foraminifera of the underlying Shekhan Formation, with similar results being recovered based on shallow benthic foramins at Chorlakki, Ganda Kas and Gali Jhagir (all dated to SB12 to SB13).

[1] The Kuldana Formation featured several different environments across its time of deposition, with the oldest layers showing fluvial conditions.

The strata overlying these sediments suggest that the environment shifted to a mix of freshwater and shallow marine habitats, which is supported by multiple lines of evidence.

It is thought that the archaeocetes that inhabited the formation at the time stuck to freshwater biomes, even if some localities were evidently closer to the coast.

[3] This mix of fresh- and saltwater habitats was eventually followed by a marine transgression that characterizes the geology of the uppermost layers of the formation.

[3] Fossiliferous sediments in and around the Ganda Kas localities were deposited in semi-arid Eocene floodplains and freshwater channels.

The most common fossil mammals are the early artiodactyl Diacodexis pakistanensis and the tapiroid perissodactyl Karagalax mamikhelensis; the presence of adapids and arctocyonids in the area indicates that it probably had an older age than the Ganda Kas fauna.