Larynx

The larynx (/ˈlærɪŋks/), commonly called the voice box, is an organ in the top of the neck involved in breathing, producing sound and protecting the trachea against food aspiration.

[2] The triangle-shaped larynx consists largely of cartilages that are attached to one another, and to surrounding structures, by muscles or by fibrous and elastic tissue components.

The cavity of the larynx extends from its triangle-shaped inlet, to the epiglottis, and to the circular outlet at the lower border of the cricoid cartilage, where it is continuous with the lumen of the trachea.

The slit-like space between the left and right vocal cords, called the rima glottidis, is the narrowest part of the larynx.

The laryngeal skeleton consists of nine cartilages: three single (epiglottic, thyroid and cricoid) and three paired (arytenoid, corniculate, and cuneiform).

Injury to one of the recurrent laryngeal nerves produces hoarseness, if both are damaged the voice may or may not be preserved, but breathing becomes difficult.

In newborn infants, the larynx is initially at the level of the C2–C3 vertebrae, and is further forward and higher relative to its position in the adult body.

The portion of the cavity of the larynx above the vocal folds is called the laryngeal vestibule; it is wide and triangular in shape, its base or anterior wall presenting, however, about its center the backward projection of the tubercle of the epiglottis.

This source sound is altered as it travels through the vocal tract, configured differently based on the position of the tongue, lips, mouth, and pharynx.

Vocal cord length and tension can be controlled by rocking the thyroid cartilage forward and backward on the cricoid cartilage (either directly by contracting the cricothyroids or indirectly by changing the vertical position of the larynx), by manipulating the tension of the muscles within the vocal cords, and by moving the arytenoids forward or backward.

The most important role of the larynx is its protective function, the prevention of foreign objects from entering the lungs by coughing and other reflexive actions.

Throat clearing is less violent than coughing, but is a similar increased respiratory effort countered by the tightening of the laryngeal musculature.

[14] Another important role of the larynx is abdominal fixation, a kind of Valsalva maneuver in which the lungs are filled with air in order to stiffen the thorax so that forces applied for lifting can be translated down to the legs.

Grunting while lifting heavy objects is the result of some air escaping through the adducted vocal cords ready for phonation.

[14] During swallowing, elevation of the posterior portion of the tongue levers (inverts) the epiglottis over the glottis' opening to prevent swallowed material from entering the larynx which leads to the lungs, and provides a path for a food or liquid bolus to "slide" into the esophagus; the hyo-laryngeal complex is also pulled upwards to assist this process.

Stimulation of the larynx by aspirated food or liquid produces a strong cough reflex to protect the lungs.

Some researchers, such as Philip Lieberman, Dennis Klatt, Bart de Boer and Kenneth Stevens using computer-modeling techniques have suggested that the species-specific human tongue allows the vocal tract (the airway above the larynx) to assume the shapes necessary to produce speech sounds that enhance the robustness of human speech.

[25] In contrast, though other species have low larynges, their tongues remain anchored in their mouths and their vocal tracts cannot produce the range of speech sounds of humans.

The ability to lower the larynx transiently in some species extends the length of their vocal tract, which as Fitch showed creates the acoustic illusion that they are larger.

Research at Haskins Laboratories in the 1960s showed that speech allows humans to achieve a vocal communication rate that exceeds the fusion frequency of the auditory system by fusing sounds together into syllables and words.

The cartilages surrounding the larynx are apparently a remnant of the original gill arches in fish, and are a common feature, but not all are always present.

While the larynx is the main sound producing organ in túngara frogs, it serves a higher significance due to its contribution to mating call, which consist of two components: 'whine' and 'chuck'.

As a result, many reptiles and amphibians are essentially voiceless; frogs use ridges in the trachea to modulate sound, while birds have a separate sound-producing organ, the syrinx.

The basic parts of the human larynx.
Vocal cords abducted and adducted
Extrinsic laryngeal muscles
Image of endoscopy
Cut through the larynx of a horse
(frontal section, posterior view)
1 hyoid bone ; 2 epiglottis ; 3 vestibular fold ; 4 vocal fold ; 5 ventricularis muscle ; 6 ventricle of larynx ; 7 vocalis muscle ; 8 thyroid cartilage ; 9 cricoid cartilage; 10 infraglottic cavity ; 11 first tracheal cartilage; 12 trachea