[1] After education at Leighton Park School in Reading South England he studied at King's College, Cambridge, from 1896 onward.

[1] He was an early Mendelian geneticist who discovered sex linkage, while writing up breeding experiment results of the Reverend G.H.

His book Heredity in the Light of Recent Research (1910), is notable for explicitly dismissing Lamarckian inheritance.

[7] He became University Lecturer in Zoology in 1914 and won the Trail Medal of Linneaean Society in 1915.

During the First World War he served as a bacteriologist to the First Eastern General Hospital, Cambridge, and later in the Friends' Ambulance Unit at Dunkirk, as he was a Quaker.