Alouatta belzebul ululata The Maranhão red-handed howler (Alouatta ululata) is an endangered species of howler monkey endemic to forests (for example Babaçu forests) in the northeastern Brazilian states of Ceará, Maranhão and Piauí.

[1] It was previously thought to be a subspecies of the red-handed howler, but unlike that species, the Maranhão red-handed howler is strongly sexually dichromatic.

[2] This New World monkey-related article is a stub.

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