[citation needed] Based on the work by Villarreal et al. 2015[2] Marchantia Dumortiera Cryptomitrium Mannia Asterella Reboulia Plagiochasma Aitchisoniella Clevea Peltolepis Athalamia Sauteria Monoclea Conocephalum Oxymitra Ricciocarpos Riccia Targionia Wiesnerella Monosolenium Cyathodium Corsinia Stephensoniella Cronisia Exormotheca Extinct complex thalloid liverworts are often represented by coalified compressions that preserve superficial morphological traits and do not allow exhaustively analysing their fine anatomy; though, in exceptional cases, fossils might preserve cell details.

Marchantites cyathodoides (Townrow) H. M. Anderson (Middle Triassic), for instance, is a Marchantia-like fossil whose detailed morphological characters (e.g., thallus with midrib, reduced air chambers, rhizoids and ventral scales) suggest a nested position within Marchantiales.

Summary tree based on the work by Flores et al. 2020:[6] Takakia ceratophylla Haplomitriopsida Pallaviciniites sandaolingensis + Pellidae Pleuroziales Metzgeriothallus sharonae + Metzgeriales Jungermanniidae Blasiidae Neohodgsoniales Sphaerocarpales Lunulariaceae Marchantites cyathodoides + Marchantites huolinhensis + Marchantia Dumortieraceae Aytoniaceae Cleveaceae Wiesnerellaceae Targioniaceae Monosoleniaceae Monocleaceae Conocephalaceae Oxymitraceae Ricciocarpos Riccia Ricciopsis + Cyathodiaceae Corsiniaceae Liverworts, possess a compelling narrative of evolutionary intricacies, particularly within the realm of complex thalloid liverworts.

The genus Hepaticites, spanning Carboniferous strata of various regions, presents a puzzling case as its affiliation with the complex thalloid liverworts dependen on its individual species.

In contrast, total-evidence dating paints a more ancient picture, tracing the complex thalloid liverworts back to the Silurian-Devonian boundary, highlighting a narrative of morphological stability across epochs.