Introduced in 1917, it benefited from significant developments of the Mark I tank (the intervening designs being small batches used for training).

The director of the Tank Supply Department, Albert Gerald Stern, first intended to fit the Mark IV with a new engine and transmission.

The Mark IV Male initially carried three Lewis machine guns – one in the cab front and one in each sponson[nb 1] – and a QF 6 pdr 6 cwt gun in each sponson, with its barrel shortened as it had been found that the longer original was apt to strike obstacles or dig into the ground.

Despite its vulnerable barrel and a tendency to overheat or foul after prolonged firing, the Lewis had the advantage that it used compact pan magazines which could hold up to 96 rounds.

A large wooden beam, reinforced with sheet metal, was stored across the top of the tank on a set of parallel rails.

The Mark IV was built by six manufacturers: Metropolitan (the majority builder), Fosters of Lincoln, Armstrong-Whitworth, Coventry Ordnance Works, William Beardmore & Company and Mirrlees, Watson & Co., with the main production being in 1917.

Crossing dry but heavily cratered terrain, many of the 60-plus Mark IVs lagged behind the infantry, but several made important contributions to the battle.

In 1940 it was restored to operational status and driven to the mainland, where its new career was allegedly brought to an early end after damaging a car.