It contains many agronomically important genera, including Arachis (peanut), Cicer (chickpea), Glycine (soybean), Medicago (alfalfa), Phaseolus (common bean), Trifolium (clover), Vicia (vetch), and Vigna (mung bean).

This clade is circumscribed by the presence of a 50 kb inverted DNA sequence in the plastome of member species and is consistently resolved in molecular phylogenies.

[3] The clade has the following ICPN-compliant, node-based definition: The most inclusive crown clade exhibiting the structural rearrangement in the plastid genome (inversion of a ~50 Kb segment in the large-single copy region with endpoints between the accD and trnK regions[10]) homologous with that found in Aldina latifolia Spruce ex Benth.

1870, Holocalyx balansae Micheli 1883, Maackia amurensis Rupr.

1917, where these taxa are extant species included in the crown clade defined by this name.