The eyes, external ears and tail are reduced to adapt to their partially subterranean lifestyle.

[citation needed] The woodland vole lives throughout the eastern United States, ranging as far as Nebraska, Kansas, Oklahoma, and Texas.

Voles prefer wooded areas with high vertical vegetative stratification but also evergreen shrubs, ground cover, and old fallen logs.

[3] Deciduous forests with moist, friable soils are suitable for burrowing and voles are most abundant in these habitats.

[2] Voles feed on both the roots and stem system and the vegetation of plants, as well as fruits, seeds, bark, subterranean fungus and insects.

[5] Voles spend most of their time underground in their burrow systems and seldom venture into the surface.

[4] Woodland voles live in family groups in burrow systems in home ranges around 14.75-17.75 in (40–45 cm).

[6] Staying in a group as a non-breeding individual is beneficial as burrow systems are major investments and a limited resource.

[9] Woodland voles create high economic loss through the damage they cause to apple orchards.