Guest stars in this episode include Neal Bledsoe, Halley Feiffer, Ryan McGinnis, Donna Murphy and Kathy Najimy.
In the episode, Betty Suarez (America Ferrera) finally has her braces removed, but only after she enters a dream sequence in which she sees what her life would be like if she had had perfect teeth.
At the same time, Justin Suarez (Mark Indelicato) tries to contact Austin (Ryan McGinnis), who he kissed in the previous episode and Daniel Meade (Eric Mabius) secures the Million Dollar Bra for a MODE cover shoot.
According to the Nielsen ratings system, the episode was watched by 4.63 million households during its original broadcast, and received a 1.7 rating/5 share among viewers in the 18–49 demographic.
At MODE, Betty is depressed that her braces have not been removed, so Marc (Michael Urie) photoshops her image into a blond haired and braceless woman on her ID.
Meanwhile, Amanda (Becki Newton) lets Tyler sleep over for the night after their dinner date due to him being jet lagged from the flight.
Her braces then set off the metal detector and when she calls out Allison's nickname, A-Bomb, she gets tackled to the ground by security and hits her head.
[2] In April 2009, series creator Silvio Horta told Michael Ausiello of Entertainment Weekly, that Betty's braces would be removed by the end of the fourth season.
"[5] Other guest stars included Halley Feiffer as Betty's friend, Allison, Ryan McGinnis as Austin and Neal Bledsoe in his ongoing role of Tyler.
[10] Tanner Stransky from Entertainment Weekly called "Million Dollar Smile" "a stunning episode" and added that it was "truly heartwarming".