Nanosaurus

Nanosaurus ("small or dwarf lizard") is an extinct genus of neornithischian dinosaur that lived about 155 to 148 million years ago, during the Late Jurassic in North America.

The taxon has a complicated taxonomic history, largely the work of Marsh and Peter M. Galton, involving the genera Laosaurus, Hallopus, Drinker, Othnielia, and Othnielosaurus, the latter three now being considered to be synonyms of Nanosaurus.

It had historically been classified as a hypsilophodont or fabrosaur, types of generalized small bipedal herbivore, but more recent research has abandoned these groupings as paraphyletic and Nanosaurus is today considered a basal member of Neornithischia.

In 1877, Marsh named two species of Nanosaurus in separate publications, based on partial remains from the Morrison Formation of Garden Park, Colorado.

Two species were named: the type species L. celer, based on parts of eleven vertebrae (YPM 1875);[5] and the "smaller" L. gracilis, originally based on a back vertebra's centrum, a caudal centrum, and part of an ulna (review by Peter Galton in 1983 finds the specimen to now consist of thirteen back and eight caudal centra, and portions of both hindlimbs).

In 1973, he and Jim Jensen described a partial skeleton (BYU ESM 163 as of Galton, 2007) missing the head, hands, and tail as Nanosaurus?

[10] Several decades later, in his 2007 study of the teeth of Morrison ornithischians, Galton concluded that the holotype femur of Othnielia rex is not diagnostic, and reassigned the BYU skeleton to Laosaurus consors, which is based on better material.

Regarding Nanosaurus agilis, Galton considered it a potentially valid basal ornithopod, and noted similarities to heterodontosaurids in the thigh bone.

[15] It was a small animal, with specimens previously assigned to Drinker and Othnielosaurus measuring 2–2.2 metres (6.6–7.2 ft) long and weighing 20–30 kilograms (44–66 lb).

[12] Heterodontosauridae Eocursor Thyreophora Lesothosaurus Agilisaurus Hexinlusaurus Yandusaurus Nanosaurus Haya Jeholosaurus Changchunsaurus Orodromeus Koreanosaurus Zephyrosaurus Yueosaurus Thescelosaurus Marginocephalia Ornithopoda Nanosaurus was one of the smaller members of the diverse Morrison Formation dinosaur fauna, diminutive in comparison to the giant sauropods.

Such dinosaurs as the theropods Ceratosaurus, Allosaurus, Ornitholestes, and Torvosaurus, the sauropods Apatosaurus, Brachiosaurus, Camarasaurus, and Diplodocus, and the ornithischians Camptosaurus, Dryosaurus, and Stegosaurus are known from the Morrison.

Holotype dentary and ilium
Type specimen of Nanosaurus agilis Marsh 1877 (YPM VP 1913) as illustrated in 1908 (left) and cast of the bone impressions after the fragmentary bone was removed (right)
Othniel Charles Marsh 's 1896 skeletal restoration of " Laosaurus " consors (now Nanosaurus ).
Skeleton with outdated skull-shape based on Hypsilophodon
Size compared to a human
Skull-less skeleton of a juvenile Nanosaurus skeleton (BYU 163)
Torvosaurus mounted as if chasing Nanosaurus , Museum of Ancient Life