[1] The northern part of the nature park begins in the Emsland near Herzlake and extends from there over the southern fringes of the Oldenburger Münsterland and Osnabrück Land in a southeast direction over the Ankum Heights to Bramsche.
From there it widens out north of Osnabrück over the Wiehen Hills eastwards and then over the gorge of Porta Westfalica, continuing further to the east as far as the Bückeburg, which lies east of the Weser and north of the Wesergebirge; the park area thus extends into the northwestern part of the Wesergebirge.
It starts east of Hörstel and runs through the Tecklenburg Land and Bad Iburg in a southeasterly direction to Bielefeld.
[2] East of Bückeburg it meets the Weser Uplands Schaumburg-Hamelin Nature Park.
Southeast of Bielefeld it runs into the Teutoburg Forest / Eggegebirge Nature Park.