In the episode, Leslie (Amy Poehler) is determined to find Ann (Rashida Jones) a date for Valentine's Day, Ben (Adam Scott) struggles with a scavenger hunt, and Chris (Rob Lowe) is depressed about his love life.
She asks the parks department to set Ann up with someone at a Valentine's Day singles mixer that they are organizing.
Meanwhile, Leslie has given Ben a Valentine's Day scavenger hunt filled with intricate riddles hidden all over Pawnee in order to discover where she wants to meet him that night, but she made it exceedingly difficult.
A grateful Ann decides to go home, but Leslie catches her applying make-up in her car.
The next day, Ron sheepishly asks Leslie to organise a riddle scavenger hunt for his birthday.
Christopher Peck of Blast Magazine said that the episode's writers "challenged themselves...while not sacrificing the consistency and rhythm of its rise-and-fall joke," and that the "'operation' was successful."