[1] However, although some phylogenetic studies have established the monophyly of the subtribes, the generic boundaries are unclear, with many genera as traditionally circumscribed being paraphyletic or even polyphyletic.
[3] A 2017 molecular phylogenetic study found that both subtribes did form clades, but did not formally recognize Habenariinae, because of missing genera and uncertainty over generic boundaries.
[3][2][4][5][6][1] As of 2017, Orchideae is divided into 6 subtribes: Brownleeinae, Pachitinae, Huttonaeinae, Orchidinae, Coryciinae, and Disinae.
[7] Once divided into the Orchidinae and Habenariinae[8] (a nonsystematic paraphyletic splitting), the Orchideae now comprise 6 subtribes.
[9][7] Along with previous molecular studies, Chase et al's 2015 analysis confirmed that Diseae was a polyphyletic grouping.