[2] The political center of the Dukagjini family was Lezhë until 1393 when it was surrendered to Venice in order to not fall under the Ottomans.
[3] The Dukagjini family appears for the first time in historical record in 1281, when their progenitor Gjin Tanushi took the title of dux.
In June 1393, the two Dukagjini brothers ceded it to Venice, which it kept until 1478 - despite later attempts by the family to retake the city.
In 1398, Little Tanush (son of Pal I), surrendered to the Ottomans, but in 1402, after their defeat at Ankara and the capture of Sultan Bayezid I, he freed himself from them.
Pal Dukagjini and his son Nikollë participated in Skanderbeg's assembly of Lezha on March 2, 1444, as vassals of Lekë Zaharia, the lord of Sati and Danja.