In a vascular plant, the stele is the central part of the root or stem[1] containing the tissues derived from the procambium.
The concept of the stele was developed in the late 19th century by French botanists P. E. L. van Tieghem and H. Doultion as a model for understanding the relationship between the shoot and root, and for discussing the evolution of vascular plant morphology.
[2] Now, at the beginning of the 21st century, plant molecular biologists are coming to understand the genetics and developmental pathways that govern tissue patterns in the stele.
[citation needed] Moreover, physiologists are examining how the anatomy (sizes and shapes) of different steles affect the function of organs.
[9] Siphonosteles often have interruptions in the vascular strand where leaves (typically megaphylls) originate (called leaf gaps).