Pterostichus is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae.

The beetles are predatory, but sometimes feed on strawberries.

They can be found under rocks and prefer slightly moist, sandy soil although preferences differ between species.

[1] These species and subgenera are included in the genus Pterostichus.

[2] Subgenus Abea Morita, 1992 Subgenus Adelopterus Reitter, 1886 Subgenus Adelosia Stephens, 1835 Subgenus Agastillus Reitter, 1892 Subgenus Anilloferonia Van Dyke, 1926 Subgenus Anomostichus Sciaky, 1995 Subgenus Aphaonus Reitter, 1887 Subgenus Argutor Dejean, 1821 Subgenus Asioplatysma Kryzhanovskij, 1968 Subgenus Badistrinus Motschulsky, 1866 Subgenus Bothriopterus Chaudoir, 1835 Subgenus Calopterus Chaudoir, 1838 Subgenus Carllindrothius Habu, 1984 Subgenus Cheporus Latreille, 1829 Subgenus Chinapterus O.Berlov, 1998 Subgenus Circinatus Sciaky, 1996 Subgenus Cophosus Dejean, 1821 Subgenus Cryobius Chaudoir, 1838 Subgenus Cylindrocharis Casey, 1918 Subgenus Eosteropus Tschitscherine, 1902 Subgenus Ethira Andrewes, 1936 Subgenus Falsargutor Kryzhanovskij, 1983 Subgenus Feronidius Jeannel, 1942 Subgenus Feronina Casey, 1918 Subgenus Fukienostichus Lafer, 1983 Subgenus Gastrosticta Casey, 1918 Subgenus Georgeballius Habu, 1984 Subgenus Gutta Wrase & J.Schmidt, 2006 Subgenus Haplomaseus Reitter, 1896 Subgenus Haptotapinus Reitter, 1886 Subgenus Huaius Tian; Huang; Chen & Ding, 2019 Subgenus Hypherpes Chaudoir, 1838 Subgenus Jedlickaia Sciaky, 1997 Subgenus Koreonialoe Park & Kwon, 1996 Subgenus Kozyrevius O.Berlov, 1998 Subgenus Lamenius Bousquet, 1999 Subgenus Lenapterus O.Berlov, 1996 Subgenus Leptoferonia Casey, 1918 Subgenus Lianoe Gozis, 1882 Subgenus Lyrothorax Chaudoir, 1838 Subgenus Megabea Sciaky, 1997 Subgenus Melanius Bonelli, 1810 Subgenus Metallophilus Chaudoir, 1838 Subgenus Micronialoe Park; Kwon & Lafer, 1996 Subgenus Monoferonia Casey, 1918 Subgenus Morphohaptoderus Tschitscherine, 1898 Subgenus Myosodus Fischer von Waldheim, 1823 Subgenus Neohaptoderus Tschitscherine, 1898 Subgenus Nialoe Tanaka, 1958 Subgenus Oreolyperus Tschitscherine, 1901 Subgenus Oreophilus Chaudoir, 1838 Subgenus Oreoplatysma Jakobson, 1907 Subgenus Orientostichus Sciaky & Allegro, 2013 Subgenus Orsonjohnsonus Hatch, 1933 Subgenus Paraferonia Casey, 1918 Subgenus Parahaptoderus Jeanne, 1969 Subgenus Paralianoe Ishida, 1958 Subgenus Parapterostichus Desbrochers des Loges, 1906 Subgenus Petrophilus Chaudoir, 1838 Subgenus Phaenoraphis Tschitscherine, 1901 Subgenus Phonias Gozis, 1886 Subgenus Platypterus Chaudoir, 1838 Subgenus Platysma Bonelli, 1810 Subgenus Plectes Fischer von Waldheim, 1822 Subgenus Pledarus Motschulsky, 1866 Subgenus Pseudethira Sciaky, 1996 Subgenus Pseudoferonina Ball, 1965 Subgenus Pseudohaptoderus Tschitscherine, 1888 Subgenus Pseudomaseus Chaudoir, 1838 Subgenus Pseudorambousekiella Schweiger, 1967 Subgenus Pseudorites Ganglbauer, 1891 Subgenus Pseudosteropus Chaudoir, 1838 Subgenus Pterostichus Bonelli, 1810 Subgenus Rambousekiella Knirsch, 1925 Subgenus Rhagadus Motschulsky, 1866 Subgenus Sinoreophilus Sciaky, 1996 Subgenus Sinosteropus Sciaky, 1994 Subgenus Sphodroferonia Kasahara & Y.Ito, 1987 Subgenus Steropanus Fairmaire, 1889 Subgenus Steropus Dejean, 1821 Subgenus Tausternus Fedorenko, 2020 Subgenus Tinautius Mateu, 1997 Subgenus Tschitscherinea Berg, 1898 Subgenus Tubuliphallus Sciaky & Allegro, 2013 Subgenus Unitrichus Sciaky, 1997 Subgenus Vietosteropus Fedorenko, 2017 Subgenus Wraseiellus Shi & Sciaky, 2013 Unassigned Subgenus Media related to Pterostichus at Wikimedia Commons