[1] The relationship of this taxon to other taxa within the division is unknown (incertae sedis), and it has not yet been placed with certainty into any class, order, or family.

The mycelium, or fungal network, can be either superficial or submerged in the substrate, presenting a coloration from yellow to brown.

[3] The spores, or conidia, are dry, ranging from subovate (egg-shaped) to obpyriform (pear-shaped), and typically have three transverse septa, though sometimes four.

The apical cell is thinner-walled and subhyaline (almost glassy), with a small conical protrusion at the tip that can produce smaller secondary spores known as phialoconidia.

[3] Quadracaea is closely related to the genera Uberispora and Physalidiopsis based on the presence of a synanamorph and rhexolytic conidial secession.

They have been found on rotten wood, branches, twigs, bark, and leaves of various trees and shrubs, as well as on decaying herbaceous materials.