The ecoregion is a strip of low forests, lakes and wetlands, running from the Baltic Sea east to the Urals.

Forest cover is typically mixed conifer and deciduous trees, with the unbroken tracts under pressure from agriculture.

[5] For aquatic habitat, the park is in the "Volga-Ural" freshwater ecoregion (WWF ID#410), a region that is characterized in general by a high number fish species but relatively few endemics.

This climate is characterized by large swings in temperature, both diurnally and seasonally, with mild summers and cold, snowy winters.

In 2012, the World Wildlife Fund initiated a project entitled "Conservation and restoration of rare bird species in large wetlands of Smolensk Lakeland National Park".

[1] The park is developed with recreational facilities, including a popular spa "Holy Well", and an ancient settlement Verzhavska.

There is a trail for children called "Fairy tales of a Russian forest", which features defense outposts and homes of folklore characters 'Baba-Yaga' and 'Leshy'.