He then completed his residency and fellowship in infectious diseases at Johns Hopkins School of Medicine.
[1] While serving in this role, he helped establish the first academic center devoted to biodefense alongside Tara O'Toole[2] and was a principal designer, author, and controller of the Operation Dark Winter[3] Later, he was also promoted from senior fellow to deputy-director of the Johns Hopkins Center for Civilian Biodefense Studies.
[1] Upon joining Pitt Med, Inglesby and O'Toole helped launch and direct the Center for Biosecurity of UPMC.
[5] In 2009, Inglesby replaced O'Toole as director and chief executive officer of the center.
[7] During the COVID-19 pandemic, Inglesby provided technical guidance to response efforts at the global, federal, state, and local level.