Togocetus

It was embedded in a phospharenite bone bed dating back to 46 – 44 million years ago, and overlying an older rock unit, the Tabligbo Group.

[3] The remains were described in 2014 by Philip D. Gingerich and Henri Cappetta, who established for it the new monotypic genus Togocetus and the new species T. traversei, dedicated to Michel Traverse.

[4] It shared many similarities with the protocetid genera Protocetus and Pappocetus, the main differences being a smaller mandibular canal, the loss of the fovea capitis femoris (hence of the ligament of head of femur) and some traits related to the molar trigonids.

[5] Later analysis of ear structure evidence originally attributed to Togocetus raised some question regarding established understanding of protocetid evolution.

Despite the fragmentary nature of the fossil record and the subtleties differentiating protocetids, T. traversei has consistently been shown to have unique features defining it, including a completely novel entoglenoid shape and several fossae otherwise not seen in contemporary cetaceans.

"Protocetid (cetacea, artiodactyla) bullae and petrosals from the Middle Eocene locality of KPOGAMÉ, Togo: New insights into the early history of cetacean hearing".