Its headwaters drain the southeasternmost portion of Villarrica National Park, including the northwestern slope of Lanín volcano.

The river flows initially to the north and from its junction with its main tributary, the Maichín River, the Trancura turns in a westerly direction until emptying into the Villarrica Lake, after having received the waters of Caburgua Lake through Carrileufú River.

The town of Curarrehue is located near its confluence with the Maichín River.

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