Tylopoda

Camelidae and numerous prehistoric families (see text) Tylopoda (meaning "calloused foot")[1] is a suborder of terrestrial herbivorous even-toed ungulates belonging to the order Artiodactyla.

It was assigned to Ruminantia by Matthew (1908); to Artiodactyla by Flower (1883) and Carroll (1988); to Neoselenodontia by Whistler and Webb (2005); and to Cetartiodactyla by Ursing et al. (2000) and by Agnarsson and May-Collado (2008).

[3][4][5] The main problem with circumscription of Tylopoda is that the extensive fossil record of camel-like mammals has not yet been thoroughly examined from a cladistic standpoint.

The first major modern and comprehensive analysis of the problem (in 2009) supported this; while some taxa traditionally considered Tylopoda could be confirmed to belong to this suborder (and a few refuted), the delimitation of this group is still very much disputed despite (or because of) an extensive fossil record.

[6] The taxa currently assigned (with some reliability) to Tylopoda are:[6] Basal and incertae sedis Superfamily Cameloidea Superfamily †Merycoidodontoidea (=Oreodontoidea) Several additional prehistoric (cet)artiodactyl taxa are sometimes assigned to the Tylopoda, but other authors consider them incertae sedis or basal lineages among the (Cet)artiodactyla or as more closely related to other artiodactyl groups like ruminants: Some studies have considered Protoceratidae closely related to Tylopoda, but others have considered them more closely related to the ruminants.

Life restoration of Agriochoerus antiquus
Life restoration of the primitive artiodactyl Diacodexis pakistanensis (foreground) being stalked by Pakicetus