In 1605 Cardinal Scipione Borghese, nephew of Pope Paul V and patron of Bernini, began turning this former vineyard into the most extensive gardens built in Rome since Antiquity.

In 1882, President Chester A. Arthur appointed William Waldorf Astor Minister to Italy, a post he held until 1885.

[2] In 2004, a colony of small Mediterranean land snails of the species Papillifera bidens was discovered living on the Borghese Balustrade.

Presumably, this species, new to the English fauna, was accidentally imported along with the balustrade in the late 19th century and managed to survive the intervening winters to the present day.

Architect Antonio Asprucci and his son Mario worked on landscaping the villa's gardens, from 1782 for over twenty years.

Camillo Borghese threw grandiose shows and popular festivals, such as a ride in an air balloon from the Piazza di Siena.

Great tit Parus major Birdlife located in the small artificial lake: Ichthyofauna: Mammals: Reptiles: Finally, among the xylophagous insects, it is worth mentioning the great capricorn beetle (Cerambyx cerdo) visible in late spring towards sunset on the tree trunks.