[4][5][6] In the male genitalia, the uncus consists of a single head bearing stiff chaetae, or it is bicapitate (Niphopyralis).

The valvae are slender and tapering towards the apex, with the costa weakly to strongly concave (e.g. in Apilocrocis novateutonialis; see [2]: 444 ).

On its mesal side, the valva sacculus is produced as a strongly sclerotised arm that usually ends dorsally in a broad, spinulose tip or a needleshaped projection, and in Aristebulea, Mimetebulea and Pseudebulea, the mediodorsal sacculus bears a medially directed process.

The signum in the corpus bursae is either absent (Mimetebulea, Niphopyralis) or present as a small to large rounded to short transverse sclerotisation.

[5][1] The placement of Niphopyralis, and with this the status of Wurthiini, was long unclear due to its unusual Limacodidae-like habitus and the lack of a proboscis.