[1] After the training of the division ended in April 1941, it was sent to East Prussia and took part in Operation Barbarossa.

[1] As part of Army Group Center, the division fought in several battles of the central sector of the Eastern Front, such as those in Białystok, Smolensk and Vyazma.

The following year it took part in the heavy fighting near Rzhev, where it remained from November 1941 to early March 1943.

[1] Due to the high losses suffered in central Belorussia, namely in Bobruisk and Baranavichy, it was reorganised in July 1944.

[1] The 129th Infantry Division was eventually destroyed in East Prussia and disbanded around February.