Accessory fruit

In other words, the flesh of the fruit develops not from the floral ovary, but from some adjacent tissue exterior to the carpel (for example, from receptacles or sepal).

[1] Accessory fruits are usually indehiscent, meaning that they do not split open to release seeds when they have reached maturity.

[11] Current research has proposed that a single class of genes may be responsible for regulating accessory fruit formation and ripening.

[12] A study using strawberries concluded that hormone signaling pathways involving gibberellic acid and auxin affect gene expression, and contribute to the initiation of accessory fruit development.

[6] Metabolic modifications in different developing accessory fruit tissues are due to the varied distributions of compounds such as triterpenoids and steroids.

A selection of accessory fruits (from left to right: pear , fig , and strawberry )
Apple section, showing seeds plus papery expression of the ovary, surrounded by tissue formed from ripening of the hypanthium.
On this strawberry, the many pips located on the surface have germinated in a phenomenon known as vivipary . The pips of the strawberry are its true fruit.
The cashew 'apple' and its attached drupe , which contains the edible seed.
Pineapple in flower