Alexander Kovalevsky

Alexander Onufrievich Kovalevsky (Russian: Алекса́ндр Ону́фриевич Ковале́вский; 7 November 1840 – 1901) was a Russian embryologist, who studied medicine at the University of Heidelberg and became professor at the University of St Petersburg.

[1][2][3][4] He was the brother of the paleontologist Vladimir Kovalevsky, and the brother-in-law of the mathematician Sofya Kovalevskaya.

[1][2][3][4] Kovalevsky discovered that tunicates are not molluscs, but that their larval stage has a notochord and pharyngeal slits, like vertebrates.

19th-century zoology thus converted embryology into an evolutionary science, connecting phylogeny with homologies between the germ layers of embryos, foreshadowing evolutionary developmental biology.

[5] He was elected on the 1st of May 1884 a Foreign Member of the Linnean Society of London.

A. Lancelet (a chordate), B. Larval tunicate , C. Adult tunicate. Kovalevsky saw that the notochord (1) and gill slit (5) are features shared by tunicates and vertebrates.
Antoine Fortuné Marion and Alexander Kovalevsky, founder and collaborator of the Annals of Natural History Museum of Marseille