Moreover, A. hulki has well-developed muscle attachments on its scapula, humerus, and ulna bones that would have allowed the forelimbs to have been held in a semi-erect stance suitable for walking over land.
Remains of A. hulki come from interbedded sandstones and marls that, based on the presence of charophyte algae, likely formed in ephemeral ponds in a large floodplain far from permanent bodies of water like lakes or rivers.
[2] A study published in 2005 had suggested that these fossils belong to several different genera of crocodylomorphs and that the original Romanian material is too fragmentary to assign to its own genus, making Allodaposuchus a nomen dubium or "dubious name".
[2] In 2013, a second species of Allodaposuchus, A. subjuniperus, was named on the basis of a skull from the late-Maastrichtian Conquès Formation, part of the Tremp Group, in the province of Huesca, Spain.
[9] In 2021, a phylogenetic analysis by Blanco disputed this result, suggesting that both A. fontisensis and A. subjuniperus belong within the genus Allodaposuchus proper, alongside the two species of Lohuecosuchus: L. megadontos and L.
[10] In 2014, A. palustris was described from a partial skull and other skeletal fragments found in Maastrichtian age sediments of the Tremp Formation in a fossil locality called Fumanya Sud in the southern Pyrenees.
A fourth species of Allodaposuchus, A. hulki, was named in 2015 and also came from the Tremp Formation, although this time in a locality called Casa Fabà.
[5] A. palustris was described by Blanco in 2021 based on fossils discovered in Late Campanian-aged fluvial deposits in Velaux-La Bastide Neuve, in Bouches-du-Rhône Department of southern France.
[12] Other studies have alternatively recovered them not as sister taxon, but rather as an evolutionary grade towards Crocodylia, with Hylaeochampsidae more basal than Allodaposuchidae.