Alvarezsauridae

They have been interpreted as myrmecophagous, adapted to prey on colonial insects such as termites, with the short arms acting as effective digging instruments to break into nests.

The sternum, for example, is elongated and deeply keeled for an enlarged pectoralis muscle, as it is in neognathous birds and volant ratites.

Martin (1997) performed a cladistic analysis but Sereno criticized it strongly, finding it flawed by incorrect codings, use of only select data, and results that did not support his conclusions.

Fossils attributed to alvarezsaurids have also been found in North and South America and Asia, and range in age from about 86 to 66 million years ago.

[9] At least one specimen of alvarezsaurid, from the species Shuvuuia deserti, preserved down-like, feathery, integumental structures covering the fossil.

Schweitzer et al. (1999) subjected these filaments to microscopic, morphological, mass spectrometric, and immunohistochemical studies and found that they consisted of beta-keratin, which is the primary protein in feathers.

[11] The discovery of Qiupanykus in association with oviraptorid eggs, indicates that the advanced alvarezsaurids may also have been specialists in nest raiding, using their robust thumb claws to crack open eggshells.

[14] Novas' 1996 description of Patagonykus, demonstrated that it was a link between the more primitive (basal) Alvarezsaurus and the more advanced (derived) Mononykus, and reinforced their monophyly.

There may be a relationship between the alvarezsaurids and the Ornithomimosauria as sister clades within either Thomas Holtz's Arctometatarsalia or Paul Sereno's Ornithomimiformes.

The discovery of Haplocheirus, which exhibits transitional features between the more derived alvarezsaurs and other maniraptorans, particularly in relation to the skull structure and development of the hand, has provided further support for that relationship.

Novas later defined the group as the most recent common ancestor of Alvarezsaurus and Mononykus plus all its descendants, though others, such as Paul Sereno, used a more inclusive definition, such as all dinosaurs closer to Shuvuuia than to modern birds.

[23]Bonapartenykus ultimus Patagonykus puertai Achillesaurus manazzonei Alvarezsaurus calvoi Ceratonykus oculatus Khulsanurus magnificus Qiupanykus zhangi Dzharaonyx eski Linhenykus monodactylus Parvicursor remotus Shuvuuia deserti Mononykus olecranus Ondogurvel alifanovi Albinykus baatar

Reconstructed skeleton of Patagonykus puertai
Size comparison of several parvicursorines: Parvicursor remotus (green), Ceratonykus oculatus (red), Shuvuuia deserti (blue), and Mononykus olecranus (violet).
Life restoration of Shuvuuia deserti