Anancus

[1][2] Recently, some authors have excluded Anancus along with other tetralophodont gomphotheres from Gomphotheriidae, and regarded them as members of Elephantoidea instead.

[16] Anancus osiris from the Pliocene of North Africa is suggested to have been a mixed feeder with a large grass intake based on microwear.

[8][6] The oldest known species of Anancus is A. perimensis, with fossils known from the Tortonian ~ 8.5 million years ago Siwalik Hills of Pakistan.

Anancus disappeared from Asia and Africa around the end of the Pliocene, approximately 2.6 million years ago.

[11] The extinction of Anancus in Africa has been attributed to competitive exclusion by elephantids, whose molar teeth were more efficient at processing grass.

Anancus arvernensis
Skull of Anancus sivalensis in side view