His skills as a geneticist drew the attention of the Red Skull, who recruited him into Hydra to aid their efforts to create super soldiers.

He finds a ready home among the Nazi Party, who see his experiments as a means to ensure the existence of the Master Race.

[8] As part of another experiment, he collects the DNA of multiple superpowered individuals and uses it to create Proto-Husks that are destroyed by Deadpool.

[10] During the "X-Men: Endangered Species" storyline, Arnim Zola was among the nine supervillain geniuses recruited by Beast to help him reverse the effects of "Decimation".

[11] In the aftermath of "Civil War", Zola joins the Red Skull in his newest attempt to kill Captain America, which succeeds.

[12] Stuck in a computer somewhere during the "Captain America: Reborn" part of the "Dark Reign" storyline, Zola is located by Norman Osborn.

Doom's machine was meant to bring him back for an unknown purpose but because of Sharon, he has become unstuck in time, causing him to relive the events of World War II.

[13] By the time Zola rebuilds the machine in Castle Doom, Sharon Carter is brought to him so that he can transfer the Red Skull's mind into her body.

There, Zola has built a massive fortress and an army of genetically altered soldiers with the intent of conquering Earth.

In the initial assault, Cap rescues a genetically engineered infant boy (Zola's son Leopold).

Eventually, Carter sacrifices herself to destroy Zola and his fortress, allowing Cap and a now reformed Jet to escape back to Earth, where only seconds have passed since his arrival in Dimension Z.

Ian finds and frees Sharon Carter, only to learn that Zola has a bomb strong enough to destroy all of New York City.

[22] A flawed copy of Arnim Zola's consciousness later joins the Shadow Council and battles the Secret Avengers, who are investigating the group's operations.

[24] An alternate universe variant of Arnim Zola from Earth-85826 appears in the Secret Wars tie-in series Hail Hydra.